C-167 “Echo” was a female Velociraptor antirrhopus masranii originally bred for InGen Security‘s I.B.R.I.S. Project. She was the second-youngest member of her subspecies to survive infancy, with two older sisters and one younger sister. She was initially planned to be a Jurassic World attraction, though InGen Security also intended to test her capabilities as a military animal. She was contained within park facilities between 2012 and 2015.
On December 22, 2015, Echo was used in a field test along with her sisters as a part of an incident taking place in the park. This incident involved the escape of a park asset, the genetically-engineered Indominus rex, which the Velociraptors were tasked with locating. Unfortunately, during the operation, Echo and her sisters rejected captivity; they briefly recognized the Indominus as an authority before rejecting it as well. In the ensuing conflict Echo was killed.
This raptor’s specimen number, C-167, comes from the mobile game Jurassic World: The Game and has not been featured in the film canon proper. Its meaning is not known. Her given name, which is used almost universally, comes from the military phonetic alphabet used by the United States Armed Forces; the letter E is referred to using “echo.” This ensures correct spelling. Echo’s trainer, Owen Grady, was a former sailor with the U.S. Navy and gave his animals (with the exception of the eldest, Blue) names from the military phonetic alphabet; Echo’s sisters have names beginning with C and D. It is unknown why the order of these letters does not correspond to the raptors’ ages. Echo’s specimen number also begins with a C while Charlie’s begins with an E; this is thought to be a mistake on the part of Ludia, Inc., the company behind the Jurassic World mobile games.
Echo was nicknamed “Elvis,” a reference to the American rock-and-roll singer Elvis Presley. They shared a signature facial expression, though in Echo this was due to a facial scar sustained during her youth.
Like her sisters, Echo was hatched in 2012 after being genetically engineered from older versions of Velociraptor antirrhopus. Echo has not been confirmed to have had any non-standard genetic modifications, making her more like the wild-type raptors than any of her sisters.
The exact date on which Echo was created and the date on which she hatched are unknown. Along with her sisters, she hatched in 2012; Blue’s hatch date is between May 15 and April 19, and Echo hatched sometime after this. She appeared to be close to her sisters in age, and was roughly eight weeks old on Day 176 of the I.B.R.I.S. Project, making her about seventeen weeks younger than the project itself.
She was the third of her kind to survive infancy, with the oldest being Blue. The next in line was Delta, with Charlie as the youngest. All four, along with their siblings who did not survive, were hatched in the Hammond Creation Laboratory after being engineered by InGen’s lead genetic biologist Dr. Henry Wu. She most likely incubated within a surrogate egg provided by another deinonychosaur owned by InGen. Upon hatching, Echo was met by her adoptive father, InGen Security animal behaviorist Owen Grady. He spent the next three years raising and caring for her.
After she hatched, Echo was fitted with a subdermal RFID tracking implant which would constantly report her location to Jurassic World’s computer network. This implant could also read her biometrics and other physiological data, making it easier to track her health. Because she was intended to spend her life within a physical paddock, her implant was probably not keyed into a particular zone of the park’s invisible fence system.
At approximately eight weeks of age, a hierarchy had developed among Echo and her sisters. Blue, the oldest, largest, strongest, and most intelligent, easily took a place at the top of this hierarchy, with her younger siblings falling in line with her. This was established by Day 176 of I.B.R.I.S., roughly twenty-five weeks into the program. Delta was Blue’s second-in-command, and Charlie was the favorite. Grady had a difficult time getting Echo to respect his authority. Echo’s loyalty to Grady was achieved only through her loyalty to Blue. Over time, Echo learned to respond to simple commands.
By 2013, Echo and her siblings entered young adulthood, which meant their lives grew more complex. They inhabited Jurassic World’s raptor research paddock, a facility including contained jungle and a training arena. Blue began challenging Grady’s authority, and he had to take measures to establish that he was still in charge. Echo, too, started to question her position in the hierarchy. She challenged Blue in a fight for dominance, and the two sisters fought violently. Blue was the victor, thanks to her superior strength and size. Echo was left with a serious laceration on the right side of her muzzle and damage to her lower jaw. Thankfully, Jurassic World’s paleoveterinary staff was able to surgically restore her to health, though her injury scarred over. This gave her a permanent sneer, causing some handlers to nickname her “Elvis” after the famous American singer.
After the fight occurred, Echo and Blue seem to have resolved their conflict and the hierarchy was maintained. Grady continued training the four raptors, teaching them more commands. By May 17, at which point Echo was about a year old, Grady came to realize that their social dynamic was more complicated than InGen originally believed. He requested that his friend Barry Sembène be recruited onto the project. His overseer Vic Hoskins agreed, and Sembène was brought on board; he became a secondary caretaker to Echo and the other raptors.
Grady performed the training exercises for Echo and the others, using a clicker device along with visual and vocal commands to teach them to perform certain actions. When they performed successfully, he would reward them with food such as beef jerky and rats. Training exercises were often unsuccessful; for example, one of the common exercises was to have the raptors chase a small pig but cease their pursuit when commanded to do so. The raptors would often let their predatory instincts override Grady’s commands, killing the pig after being ordered not to.
When the raptors were not in training, Sembène tended to their health. While the raptors tested their caretakers and frequently made escape attempts, they still maintained a sense of loyalty. They learned to recognize and respond to at least forty distinct commands. Some exercises, such as the aforementioned pig chase, were a struggle even as of 2015; however, Echo and her sisters were recorded as responding appropriately to commands 73% of the time.
This trust was not present with all of the human members of IBRIS. Many of the ACU personnel hired by Vic Hoskins were prepared at any time to use nonlethal weapons to subdue the raptors, and at least one paddock worker was severely mauled by a raptor in 2015. Grady himself was wary of the raptors and kept a close watch on all four of them to prevent escapes or attacks on any more workers. Of all the raptors, Echo seemed to be the most ready to attack a handler.
There was a brief incident during the night of December 19, 2015 when three campers from Camp Cretaceous trespassed into the raptor paddock. Echo was called by Blue to help confront Kenji Kon, who was separated from them by a gate. This gate was accidentally opened, and fellow camper Darius Bowman entered the paddock to help him. The two teenagers were rescued by the intervention of camp counselors Roxie and Dave. Floodlights blinded the raptors, and meat was delivered to distract them.
On December 22, 2015, when Echo was three years old, Blue successfully responded to Grady’s stop command during a pig chase exercise. Delta, Echo, and Charlie followed suit, deferring to Blue’s authority. This was witnessed by Vic Hoskins and other InGen personnel. Shortly after the exercise, the pig used in the chase escaped back into the paddock, and a recently-hired handler named Colby Boothman-Shepard attempted to recover the animal. While he did snare it successfully, it was grabbed by Echo while still in the snare; this caused Boothman-Shepard to be dragged off the catwalk and into the paddock. Grady entered the paddock to rescue his employee, confronting Blue while convincing ACU to stand down. Echo was feeding on the pig while this happened, and by the time she finished, Grady had gotten Boothman-Shepard out of the paddock and escaped himself. Echo came over to inspect the aftermath, her sisters having charged to kill Grady but narrowly missed him as the gate closed; Echo took the opportunity to snarl at Boothman-Shepard from inside the paddock, but the thick titanium bars kept them separated for now.
That night, Echo and her sisters were commandeered by InGen Security members ranking above the usual ACU staff. They were led into a hide-and-seek exercise, which they had performed numerous times with Grady. Echo and her sisters were presented with a piece of meat belonging to a species they had never encountered before, instructed by Grady to track down the source. They were released into the world outside for the first time, and Echo followed Blue in joyfully sprinting through the jungle on her first real hunt. Grady, Sembène, and InGen Security members followed.
When they neared the source of the scent, they slowed and stopped. An unknown animal was heard vocalizing in the jungle nearby, and Echo responded to it with a social call. Emerging from the jungle, the enormous Indominus rex revealed itself to the raptors. Blue and the new creature sized one another up, each vocalizing to try and establish dominance. During the interaction, Echo began to realize that the Indominus was a more powerful leader than Grady, and that its strength could give her pack the one thing Grady would not: the freedom to live in the wild. Echo observed Blue’s deliberation and followed suit.
InGen Security opened fire on the Indominus, frightening Echo and the other raptors into the jungle and driving the larger creature away. Now viewing the humans as a threat, Echo and the others set up ambushes to kill off the InGen soldiers one by one. During the ensuing firefight, Charlie was killed. This sent Blue into a rage, and she now led Echo and Delta in an attack on Grady himself. The chase led back to the paddock, and they began to pursue MVU-12. Blue led them to attack MVU-12 as it sped to the southwest; Blue attacked its driver Claire Dearing from the side during the chase, but was struck against a tree and fell out.
Echo and Delta coordinated to continue the chase, aiming for the still-open doors in the back of the MVU. Within were two young humans, Zach and Gray Mitchell, who fended off their attackers by throwing medical supplies and utilizing a shock prod. Shortly thereafter, the chase was called off as the Indominus summoned the three remaining raptors to reconvene in the forest.
They made their way to Main Street ahead of the Indominus, most likely following the scent of the vehicles Grady had fled with. Delta went ahead to track him through the Innovation Center, and Echo joined Blue in setting up an ambush outside. While Blue was going to stop Grady head-on, Echo was waiting in the sidelines to sabotage any attempt at escape that Grady might make. Delta flushed out Grady and his human companions, and the ambush worked as intended; with Echo cutting off the easiest escape route and Delta moving in to her opposite side, the humans had no chance at getting away. Echo and Delta awaited Blue’s kill command.
However, Grady managed to calm Blue. While the Indominus promised her freedom as her alpha, she did not provide familial love or mutual respect in the way that Grady had done for Blue’s entire life. Grady removed her harness, promising her freedom. Despite Charlie’s death, Blue calmed down and felt loyal to Grady as she had before. Although they did not understand Blue’s decision, Echo and Delta obeyed their sister.
The Indominus caught up with the raptors, commanding them to kill. Blue refused, which earned her a devastating slap from the larger animal. She was thrown into a storefront, stunned and unable to move; she appeared dead. Echo and Delta, complying with Blue’s final act as their leader, stayed loyal to Grady and attacked the Indominus. Despite delivering savage bites and slashes, their fury was not enough to bring the creature down even with help from Grady’s rifle. Its tough skin was more than their natural armaments could penetrate. Delta was tossed into Winston’s Steakhouse, landing on the gas ovens and inadvertently triggering a brief but fatal gas explosion. Echo put up a valiant attempt to continue the fight, but lasted only moments longer than Delta; she was grabbed by the Indominus and sustained crushing internal injuries from its massive jaws. Her body was tossed aside, and she died of her wounds.
Most likely, no efforts to remove Echo’s remains from Isla Nublar were made, as the park was evacuated and abandoned shortly thereafter. Her body was the most intact of her fallen sisters, and she was probably scavenged by newly-freed carnivores in the following days. Echo’s skeletal remains may have remained in the maintenance alley where she landed unless they were moved by animals or natural events.
Despite appearances, Echo was survived by one biological family member: Blue, whose injuries were not fatal, and who had been left to watch helplessly as her family was slaughtered before her eyes. She recovered enough to spring back into battle and, with the unexpected help of a freshly-released Tyrannosaurus, drive the Indominus toward the Jurassic World Lagoon where it was snatched by the park’s resident Mosasaurus and torn in half underwater.
Grady, too, survived this incident thanks to Blue’s intervention. Jurassic World was closed permanently, and Grady left the island to begin a new life. He and Blue were eventually, briefly, brought back together three years later; Grady unwillingly aided in relocating Blue to the North American mainland as Isla Nublar was destroyed by a volcanic eruption. Grady’s reluctant participation in this mission was inspired by his memories of the I.B.R.I.S. Project, including memories of Echo when she was young.
Echo was among the fiercest of her sisters, favoring offense over defense in a fight. She made excellent use of her teeth and claws to savage prey items, preferring to quickly subdue prey with a sudden ambush attack. Echo’s techniques were classic tactics used by generations of InGen Velociraptors beginning with the first ever cloned; her go-to strategy was an unexpected tackle from the side, catching the prey unaware and striking before it could react. She did not have as many genetic modifications as her sisters, so her behavior could be considered more akin to that of her ancestors who first inhabited Isla Nublar and Isla Sorna in the early 1990s.
Before her death, Echo also fought against humans and an Indominus rex. In both cases she worked alongside her sisters, letting Blue or Delta take point while she attacked from an unexpected direction. This subterfuge made her attacks difficult to predict and let her cut off means of escape. Sabotage in this manner was Echo’s area of expertise. She also did well in simple pursuit hunts, though she truly excelled when she had the element of surprise. Echo’s excellent senses gave her an edge against most animals, especially in the dark. Her I.B.R.I.S. training made her deadlier than normal in fights against humans, since she understood their behaviors better than other creatures; during the 2015 incident she killed several InGen personnel via ambush attacks in cooperation with her sisters.
Velociraptor lives in small groups of family or friends, and is social by nature. Echo’s impulsive and independent nature, however, could sometimes be problematic. During her youth, she challenged her sister Blue for the beta position in their pack (the alpha position being maintained by trainer Owen Grady); despite Blue’s superior strength and intelligence, Echo fought her anyway. The fight turned brutal, with Echo sustaining serious wounds to her jaw that required veterinary care to rehabilitate. After the fight, though, Blue and Echo seem to have made up; there were no further reports of violence between them. Echo’s interspecies social skills were also a mixed bag. She was the one to respond to the call of the Indominus, bringing the creature in to negotiate with Blue for authority, but Echo also was hard for her handlers to keep under control and she was a major threat to the humans she worked with. In that case, only Blue’s authority and Grady’s watchful eye kept her from attacking her handlers.
Physical strength and durability
Echo was as speedy and athletic as any of her kind, easily keeping pace with her sisters (even the notably powerful Blue). As described above, she was a competent brawler and skilled on the hunt, making good use of her lethal claws and teeth in a fight. Her top speed was around fifty miles per hour, and her endurance was impressive as well. Armed with a strong sense of smell and low-light vision, Echo was a capable predator despite her life in captivity, and proved able to hold her own during the brief time she spent in the wild.
She did not survive her fight with the Indominus rex, though she could overpower smaller enemies with ease. Against this massive creature, she did not really stand a chance, but survived the fight for a respectable amount of time, using all her physical attributes to gain an advantage. Her agility and speed were her main weapons, since she could maneuver better than the Indominus and move quickly enough to stay unpredictable. She attacked from the sides, clinging to the bigger animal’s back where its clawed hands and crushing jaws could not reach. When the counterattacks threatened her, she disengaged and tried again from another angle. Only after Delta was fatally wounded did Echo misstep; she suffered just one hit from the Indominus during the fight, but it was enough to kill her.
In addition to the system of communication that her species normally uses, which some scientists such as Dr. Alan Grant consider to be a language, Echo had a basic understanding of some English words and phrases as well as symbolic gestures. In total, Echo comprehended over forty distinct instructions and as of 2015 responded correctly 73% of the time. Some of the incorrect responses are known to have resulted from her deliberately ignoring commands, so her actual rate of comprehension was higher than the 73% value.
Echo also had a limited ability to understand other species, namely the Indominus she encountered during the 2015 incident. She was the one who first responded to its call, though she left most of the communication up to Blue. She efficiently understood the larger theropod’s intents and instructions. Both vocal and body language were used between the two species to facilitate communication.
On social structure
Among all of her sisters, Echo was the only one to question Blue’s leadership at any point, considering herself more fit for the role. Blue’s leadership style was empathic and strategic, in contrast to Echo’s results-first and headstrong approach. Being physically beaten by Blue in a fight seems to have tempered Echo’s ambition, and from that point on she accepted (if begrudgingly) her position in the hierarchy. She never fully accepted Owen Grady as her alpha, only obeying him out of respect for Blue; when the possibility of finding a new leader arose, Echo accepted it readily.
The Indominus seemed to be an ideal alpha at first; this creature was not only massive and powerful, but intelligent, and also utterly ruthless. Echo respected this, and also recognized the Indominus as being something like herself, better suited to be a raptor alpha than a human could be. She followed it without question after Blue accepted it as their leader. In the end, though, Blue rejected the Indominus because it did not truly care about the raptor family at all. Echo may not have had as much a sense of family loyalty as Blue did, but the Indominus lacked it entirely. The Indominus also lacked real ambition, having no distinct goal other than to cause havoc and slaughter creatures. While Echo was a savage fighter, she attacked with reason and purpose. Blue’s apparent death at the hand of the Indominus did nothing to dissuade Echo from turning against the creature, and she remained loyal to Blue’s final command until her dying breath.
InGen Security animal behaviorist Owen Grady was the first family member Echo met. He was the first creature she saw when she hatched in 2012, so she imprinted on him as a parent. From then on, he fulfilled the role of father figure. Grady had been hired by InGen to be lead researcher on the I.B.R.I.S. Project, which sought to research the animal intelligence of deinonychosaurs to better human interactions with them. However, Echo quickly began to question Grady’s right to be her leader, instead deferring to Blue.
Echo followed Blue’s lead in challenging Grady’s authority as they grew older. Blue, of course, was the strongest of them all and a better hunter than Grady. Echo did fight Blue for dominance at one point, but this subsided after she lost; Grady did not get any such respite and was always forced to defend his authority. Blue led Echo and the others in escape attempts, which Grady repeatedly had to thwart.
On December 22, 2015, Echo successfully passed a training exercise in which she was meant to resist her urge to kill a pig when Grady commanded her to stop, but when the pig was accidentally let back into the paddock, Echo recognized training was over and killed the animal. A handler was attempting to retrieve the pig when the attack occurred and was dragged into the paddock. Grady was forced to enter in order to protect the employee, and while Echo was busily eating the pig, her sisters nearly killed Grady.
That same day, Grady took Echo and her sisters in their first out-of-paddock hunt. He did not lead, instead letting the raptors chase down their target using their superior sense of smell. When they located the target, the Indominus rex, the raptors established a form of communication with it and quickly realized that this animal met all the alpha criteria: it was huge and powerful, intelligent and confident, and armed with impressive hunting implements. Grady, by comparison, was an inferior alpha. Turning on Grady was now an option on the table, and the decision was made for them when InGen Security opened fire. Echo attacked Grady at Blue’s orders after Charlie was killed in the firefight.
Echo pursued Grady and his allies to the Innovation Center, where she waited in ambush to cut off his route of escape. Delta flushed Grady out of hiding, and Echo ensured to stop him from getting away. However, Blue had cooled her head by that time, and Grady was able to calm her down; now Blue refused to kill Grady, and Echo obeyed. Even when Blue appeared to have been killed, Echo stayed true to Blue’s last order and defended Grady until her death.
Echo’s oldest sister, Blue, was the dominant member of their pack not just because of her age but because of her size, strength, and intelligence. She had another trait that made her an effective leader, and it was one Echo did not fully understand: she was empathic, able to understand what her sisters were thinking and feeling. Blue strongly favored loyalty, while Echo favored independence. They also differed in strategy; Blue would think through a battle plan, whereas Echo was impulsive and hotheaded. Their differences led to violence when they were young, as Echo eventually chose to challenge Blue for the leader position. The fight ended with Echo sustaining serious wounds to her lower jaw and snout, but being physically beaten into submission seems to have kept her ambition in check. No further fights between the two sisters were reported.
On the hunt, Echo acted as Blue’s saboteur, setting up ambushes and preventing prey or enemies from escaping. Echo’s rapid and aggressive attacks made her an unpredictable fighter, lending Blue’s pack the advantage of surprise. This was put to lethal effect during the 2015 incident. Echo managed to kill several InGen mercenaries during the conflict, using her knowledge of human behavior patterns from I.B.R.I.S. to outmaneuver her enemies. She even helped Blue corner Owen Grady himself, though Blue changed her mind about killing him. Echo was not as loyal an animal as Blue, but she did have a sense of family, and respected Blue’s decision. She fought fiercely to avenge Blue when it appeared the Indominus had killed her, dying in the effort.
Echo and Delta seem to have been close in age, with Delta being the next oldest after Echo. Delta was also the sister with whom Echo had the most in common; they appear to have been quite close. Both of them shared a sense of strong independence and a love of the hunt. Delta’s superior eyesight and single-minded focus made her an effective killing machine, which Echo respected.
The main difference between them was that while Echo was an impulsive raptor, Delta was more levelheaded and mature. Delta remained neutral in the conflict between Blue and Echo when they were younger, and never challenged Blue; instead she understood Blue’s skill as leader and respected it. Delta also had a similar respect for Echo’s skills, knowing how to direct her sister’s ferocity to great effect. This made Echo and Delta a powerful team. Often they could coordinate their efforts with nary a sound between them; body language was all they needed.
During the 2015 incident, their coordination was put on full display against InGen soldiers, with a body count racking up between the two sisters. With Delta selecting a target, her unbreakable focus and Echo’s unpredictability proved an impossible combination for InGen to beat. They later fought together against the Indominus rex when it appeared they were the only surviving raptors; they stayed together until their final moments, Echo being killed moments after witnessing Delta’s shockingly sudden death. Delta’s support seems to have been so critical to Echo’s sabotaging attacks that she could not last long against such a foe without her sister by her side.
Echo’s younger sister, Charlie, was the baby of the pack and most loyal to Blue out of all of them. While their age difference was not large, the difference in maturity between Echo and Charlie was as distinct as that between Echo and Delta. Even while Echo was hotheaded, Charlie was the most likely to give in to her instincts. This made her aggressive, a trait which Echo could appreciate, but her youthful exuberance could get the better of her. She had a fairly poor sense of spacial awareness and, due to their respective positions during hunts, was known to sometimes strike Echo in the face with her tail without realizing it. Having been thoroughly beaten for the leader position by Blue, there was little Echo could do to put her younger sister in line, so she tolerated Charlie’s oblivious hyperactivity. On the hunt, though, Charlie’s energy could be directed at their prey or a foe, making her a useful ally to Echo. Between the two of them, any enemy would be quickly overwhelmed and tired out.
Unfortunately, during the 2015 incident Charlie was among the first victims. She was killed in a firefight with InGen Security shortly after the sisters rejected Grady and began taking orders from the Indominus rex. Blue’s attack against Grady began when Charlie died, so a large portion of this struggle was an act of vengeance. By the end, though, Blue had cooled off emotionally and recognized that Charlie’s death was not due to Grady but due to the poor leadership of the Indominus. Echo seemingly concurred with this realization, and Charlie’s death was taken off Grady’s shoulders.
When Echo was roughly a year old, her family grew by one human. Owen Grady’s old friend Barry Sembéne was brought to Isla Nublar to join the I.B.R.I.S. Project as an animal caretaker, helping to manage the raptors as their social behavior was more complex than anticipated. While Grady was the primary trainer, Sembéne helped to keep the animals healthy and happy by providing close personal care to them.
Although Sembéne tended to the raptors closely for the next two years, he maintained a sense of caution about them. He was well aware of their power and skill on the hunt and hardly trusted Grady himself to be in the paddock with them. Sembéne expressed doubts that the raptors would ever be trustworthy enough to be let out of the paddock, believing that they would take any opportunity to reject human leadership and make their own way. His assessment was not inaccurate; when the raptors were presented with the Indominus in the wild, they accepted it as their new alpha in place of Grady. This led to a firefight with InGen Security, in which Charlie died.
Blue attacked Sembéne after Charlie’s death, and Echo was drawn away to other parts of the conflict. Sembéne and Echo never saw each other again; the raptors pursued Grady toward the park attractions, and Sembéne was left to make his way back to civilization alone. By the time he arrived, the conflict had already been resolved with Echo among the deceased.
Dr. Henry Wu
Like all of InGen’s dinosaurs, Echo’s genome was constructed by geneticist Dr. Henry Wu. Echo was engineered to be more suitable for the I.B.R.I.S. Project, since the older raptor varieties had failed due to unpredictable aggression. She seems to have had the fewest modifications of all her sisters, most similar to the original raptors.
Despite being Echo’s creator, Dr. Wu does not appear to have been present for much of her life. After she graduated from the Hammond Creation Lab’s hatchery, she likely did not see him often at all, since he rarely left the lab. However, Wu did also create the Indominus, which led to the end of Echo’s life in 2015.
Vic Hoskins and InGen Security
I.B.R.I.S. was an InGen Security project from its inception, with the planning stages occurring well before Echo or her sisters were even a concept. She was hatched into a program overseen by Security’s leading mind Vic Hoskins, whose aims for the project were to determine whether genetically-modified Velociraptors could be used as military animals in the same manner as dogs and other modern species. This aspect of I.B.R.I.S. was its most controversial, with some Security employees such as Grady and Sembéne strongly opposing it. Hoskins began pressing for a field test in 2013 when it was determined that the raptors’ social behavior was more complex than originally anticipated.
Although Hoskins was the project overseer, he was not frequently at the paddock. Instead, he received updates from his employees, particularly Grady and Sembéne, but also the ACU troopers employed to watch the paddock. The troopers were tasked with keeping a close eye on Echo and her sisters and acting to prevent escapes. Other staff at the paddock, including handlers such as Colby Boothman-Shepard, may have been employed by InGen Security. Two handlers including Boothman-Shepard were threatened by Echo and her sisters in 2015, with the unnamed handler being wounded so badly that they could no longer continue working at the paddock. It is not known if this handler survived the injuries. Echo threatened Boothman-Shepard during an incident in which he was dragged into the paddock, but by the time she got around to approaching him, he had been rescued from the paddock already.
During the night of December 22, 2015, Hoskins brought a number of InGen Security contractors and soldiers to Isla Nublar and commandeered I.B.R.I.S. following Simon Masrani’s death. These members of Security were not part of Jurassic World’s staff, so Echo was unfamiliar with them. InGen Security accompanied Echo and her family on their first out-of-paddock hunt, which led to her encountering the Indominus. This creature convinced Blue to abandon Grady, and Echo concurred. Moments later, the Security personnel were ordered remotely by Hoskins to open fire on the Indominus. The raptors initially fled from the gunfire but then turned on Security, using ambush tactics to pick them off one by one. Charlie was killed in the fight.
Echo did not see Hoskins or any InGen Security members again after this. Delta, while driving Grady and his allies into an ambush involving Echo and Blue, had killed Hoskins; other Security staff were busily evacuating the island. Echo was killed before any of them arrived to Main Street.
Tourists at Jurassic World were unable to see Echo or her sisters in person, since they were classified for I.B.R.I.S. research and not accessible to the general public. Her only direct interaction with visitors to Jurassic World was with two members of Camp Cretaceous, Kenji Kon and Darius Bowman, who trespassed into the raptor paddock on December 19, 2015. Echo was summoned by Blue to attack the two teenagers, but they were stopped by the park rangers along with Camp Cretaceous counselors Dave and Roxie.
On her first out-of-paddock hunt, Echo was tasked with tracking down an unknown animal by scent using a piece of its flesh. She followed Blue on a breakneck hunt through the island’s northern jungle, slowing down only when she smelled the creature nearby. They heard its vocalization, which sounded surprisingly familiar, and Echo responded to it. The animal revealed herself: an enormous theropod the likes of which she would never have seen before, the Indominus rex. This animal was able to understand the raptors’ vocalizations and respond in turn. Blue asserted her status as the pack’s beta as the dinosaurs sized one another up.
The Indominus appeared, to Echo and her sisters, to be everything a raptor would look for in an alpha and leader. She was powerful, much more so than any human or even a raptor, with deadly claws and teeth. She was also intelligent and confident, traits that are highly valued in raptor alphas. There was also one other advantage the Indominus could provide as an alpha: the freedom to live outside of the paddock walls, no longer confined or restricted. Blue rejected Grady in favor of this new and alluring leader, and Echo concurred.
With the Indominus as her alpha, Echo engaged in a fight against InGen Security which led to Charlie’s death. This enraged the raptors, who blamed the humans rather than the Indominus at first. Echo chased down an MVU as it fled from the scene, only giving up the chase when the Indominus summoned them all to reconvene in the jungle. They made their way toward their new leader’s original destination: the hotel, where all the park visitors were packed waiting to be evacuated. This worked in favor of Echo, since Grady and his friends were at the Innovation Center just across the Lagoon from the hotel. She and her sisters went ahead of the Indominus to find Grady.
They tracked him to the Innovation Center. Delta went inside to flush him out, with Echo and Blue waiting to ambush him when he was chased outside. When they had him surrounded, though, Blue hesitated and did not give the kill order. She reconsidered her decision to reject Grady in favor of the Indominus. Echo, by now, had calmed down enough to think more rationally as well: the Indominus had led them into a fight where Charlie had died almost immediately, and Grady had kept them safe for their entire lives until this point. The Indominus, despite all her strengths, lacked any motive beyond plain slaughter. She also had no sense of loyalty, driving the raptors to fight on even if it cost their lives.
When their new alpha caught up with them, she gave Blue a kill order, which Blue refused. For her defiance, Blue was given an apparently fatal slap, and Echo and Delta were given the opportunity to reconsider. But now, with the blood of their beloved sister on the claws of the Indominus, they would never respect her again. Echo and Delta stayed true to Blue’s last order to them, fighting the Indominus until their final breaths. Delta was killed in the fight, and moments later, Echo made a fatal misstep and was caught in the hybrid’s jaws. Echo was slain, but her valiant fight had given the humans enough time to take action. Blue, who had survived her attack, regained consciousness and avenged her fallen sisters by helping to drive the Indominus to her death in the Jurassic World Lagoon.
Echo’s relationship with Jurassic World’s Senior Assets Manager Claire Dearing was mostly one-sided, since Dearing mostly supervised all of the park’s animals from the control room and other operations facilities. Echo, as a specimen in a high-profile InGen Security project, would have been of particular importance. However, Dearing never actually saw Echo or her sisters in person until the 2015 incident, during which the Head of Security Vic Hoskins appropriated the I.B.R.I.S. raptors for a field test. Dearing was not involved with the exercise, but was affected by its aftermath.
After rejecting Owen Grady as her alpha and retaliating against him for Charlie’s death, the high-speed chase took Echo to the access road where Dearing was attempting to drive her nephews to safety in a Mobile Veterinary Unit. During the pursuit, Echo attacked the MVU, following Delta in an attack on the rear doors. She was unable to stop the vehicle, and before she could make another attack, she was summoned away by her new alpha. The chase then became an ambush hunt, leading to Main Street.
Although Dearing was put in the path of Echo’s attacks, she was not the main target. Instead, she was in danger through association with Grady. Echo set up an ambush for Grady outside the Innovation Center, and since Dearing was accompanying him, she and her nephews were also caught in the ambush. Although the setup was flawless, Blue had her stand down rather than make the kill. Blue had realized that the Indominus was unfit to be their alpha and called off the attack. Dearing’s life was spared as a result. Echo died in the ensuing fight, but her assault on the Indominus gave Dearing enough cover to put a new plan into action.
Other InGen staff
While access to the raptor research paddock was restricted to certain personnel, many members of InGen were at least aware of Echo’s existence. She was probably researched by geneticists due to her unique genomic constitution, and was cared for by paleoveterinarians and other animal handler staff members. The I.B.R.I.S. Project involved various other researchers, and had been authorized by Masrani Global Corporation‘s CEO Simon Masrani; he hoped that the raptors could be integrated into a park environment if they were properly understood. Masrani opposed Vic Hoskins’s plan to market the raptors as military animals.
Zach and Gray Mitchell
Echo was briefly involved with the Mitchell brothers‘ misadventures in Jurassic World during the 2015 incident. They accompanied their aunt Claire Dearing and her romantic partner Owen Grady to the raptor paddock on the night of December 22, though they were not intended to be a part of the action. Instead, they were there to ensure that Dearing could keep watch over them personally. After the Indominus seduced the raptors away from Grady, a high-speed chase down the service road between the paddock and Sector 3 ensued. The Mitchells were on board an MVU driven by Dearing and were threatened by Blue and her sisters as they pursued the vehicle.
The Mitchells were also present at the Innovation Center when Echo set up an ambush for Grady, though they were not the main target. As with Dearing, they were threatened only because of their association with Grady, and were spared from further violence when Blue and Grady made amends. Echo was killed in the fight that followed, but she kept the Indominus occupied. While watching Echo and Delta, Gray realized that the raptors were not powerful enough to win the fight, which inadvertently inspired Dearing to enlist the help of another, bigger animal. Sadly this came too late to save Echo.
Echo is portrayed through a combination of motion-capture animation, computer-generated imagery, and animatronics. Echo’s motion capture in Jurassic World was performed by an uncredited ILM employee. Like most of the characters in the sequel trilogy, she is not based on any particular character from Michael Crichton‘s novels.