Stegoceratops (S/F)

STegoceratops_jw
Digital rendering of Stegoceratops on Dr. Henry Wu’s laboratory computer, as of 12/22/2015

Stegoceratops is a transgenic hybrid genus of genasaurian dinosaur engineered by Dr. Henry Wu of International Genetic Technologies sometime prior to December 2015. Its name translates to “roofed horned face,” and appears to be a portmanteau of Stegosaurus and Triceratops. Its specific epithet is thus far undisclosed. This was one of several hybrid genera conceptualized by Dr. Wu, but there is currently no evidence that any specimens were ever created.

It is not known if this species has been assigned a family, but it cannot be reliably placed on the tree of life since it is a combination of many different organisms. Usually it is considered to be a genasaurian, since this would be inclusive of the two main species used to create its genome. However, it could also be more broadly classified as a saurian (including its snake genes), a eugnathostomatan (including its flatfish genes), a nephrozoan (including its cuttlefish and arthropod genes), or most generally a parahoxazoan (including its jellyfish genes).

Digital design of the Stegoceratops and presumed constituent organisms (created by Rudy Vessup http://www.rudyvessup.com/)

Like the Indominus rex, Stegoceratops was a highly advanced hybrid with its genome constructed using the DNA of various other organisms. Its sourced species are mostly unknown at this time, but there is evidence for the following:

  • Stegosaurus stenops: This was one of two species which were used primarily to engineer the animal’s body. Features derived from Stegosaurus include a staggered double row of dorsal plates and a thagomizer of four spikes. According to the Ludia-sourced Jurassic World Hybrids children’s book, Stegosaurus was the base genome used as a template for this animal, but there is no evidence to suggest this in the films. In some versions of the animal, smaller paired rows of plates can be seen adjacent to the main, larger paired rows.
  • Triceratops horridus: This was one of two species which were used primarily to engineer the animal’s body. Features derived from Triceratops include the overall body shape of the hybrid, particularly the skull. Some renditions of the animal depict it with curved, rather than mostly straight, horns unlike those of Triceratops, but there is currently no evidence to confirm whether any other ceratopsian species were sourced to influence its anatomy. The version of the animal in the game Jurassic World Evolution depict it with extremely long horns similar to those seen on Torosaurus, which has been interpreted as a more mature form of Triceratops by some scientists including Dr. Laura Sorkin. This version of the animal also has a small pair of smaller horns located adjacent to the large ones, as well as numerous hornlets on its frill. A few versions depict the animal with a nasal horn, but most feature a nasal boss instead.
  • Dynastinae: At least one as-of-yet unidentified species of rhinoceros beetle is depicted on a computer screen alongside Stegoceratops in an unused render of Wu’s private laboratory computer system. It is affixed with the caption “Exo-Skeletal Armor,” suggesting that elements from the beetle’s durable chitinous exoskeleton would have been incorporated into the biology of Stegoceratops. A slightly different species is also depicted on a different render, on which Stegoceratops is absent; this second variety is the one which appears on the screen in the film itself.
  • Serpentes (Crotalinae?): An unidentified species of snake appears next to Stegoceratops on Dr. Wu’s private laboratory computer. Indominus rex, which was also designed by Dr. Wu, incorporates Crotalinae genes into its genome, which gave it the ability to see in infrared; it is not known if this was the same genetic source used for Stegoceratops.
  • Sepiida: An unidentified species of cuttlefish can be seen next to computer renders of Stegoceratops. The label reads “Camouflage,” indicating that genes were sourced from the cuttlefish to facilitate the development of chromatophores. It is not known if this trait in Indominus was intentional, as Dr. Wu claimed that the cuttlefish’s DNA was sourced to encourage the survival of an expedited growth rate. It does appear to have been intentional in Stegoceratops.
  • Pleuronectiformes: An unidentified species of flatfish can be seen depicted near a computer render of Stegoceratops with a label reading “H2O – Oxygen Extraction,” indicating that genes from the flatfish may have been used to give this hybrid the ability to extract oxygen from water. This would enable it to survive longer when submerged, though the renders do not depict true gills anywhere on its body.
  • Pelagiidae: What appears to be a pelagiid jellyfish appears as a computer rendering next to that of a Stegoceratops, with the label “Bio-Luminescence.” The jellyfish Pelagia noctiluca, or mauve stinger, is known to exhibit bioluminescence, but the animal depicted in the render more closely resembles a member of the genus Chrysaora. Proteins from jellyfish are often used in genetic engineering to make organisms glow under ultraviolet light; however, the genes sourced from this pelagiid are intended for true bioluminescence, not the biofluorescence that is often utilized in genetics.
  • Unknown Invertebrate: An unidentified invertebrate organism can be seen on the renders of Stegoceratops with a caption that appears to read “Luminescence,” presumably referring to bioluminescence in the manner of Pelagiidae described above. The identity of this organism has not yet been determined.
  • Danainae: An unidentified butterfly which appears to be a danaine (for example, the monarch Danaus plexippus) appears on an unused laboratory computer screen render which also features Stegoceratops. However, they do not appear simultaneously, making it ambiguous whether Danainae DNA was incorporated into the hybrid genome. Both the Danainae and Stegoceratops versions of this same render have text reading “Increased Vision Spliced Extractioned Organic Code,” implying that if Danainae was utilized in Stegoceratops, it may have been to improve its vision abilities. Butterflies can see ultraviolet light, a trait which the hybrid animal may have had.
Design of Dr. Wu’s personal lab computer display. A nearly-identical piece shows a Stegoceratops in the main cell. Design from http://www.rudyvessup.com/

The Stegoceratops, along with other hybrids including Indominus rex, would have been designed sometime during or after April 4, 2008 and prior to December 22, 2015.

Description

Compared to the best-known hybrid genus, Indominus, the constituent parts of the Stegoceratops are plainly visible in its at-a-glance anatomy. The basic body shape is essentially the same as its ceratopsian parts, resembling a Triceratops in most ways. The body is slightly narrower, however, and the legs somewhat longer. The front feet have five toes, while the rear feet have three toes; all four of its feet are relatively small compared to its body size.

Computer display of Stegoceratops and genomic data, featuring a focus on cuttlefish DNA.

The low-hanging tail of the Stegoceratops is considerably shorter than that of Stegosaurus, but like its stegosaur component, it does have a thagomizer. The thagomizer consists of four conical bony spikes which extend outward and curve upward. Like the tail, the thagomizer itself is smaller in size than those known in Stegosaurus. The other stegosaurian feature of this animal is the dual row of roughly triangular dorsal plates. In the render shown on Dr. Wu’s computer terminal, it has fourteen plates in a staggered configuration; mobile games and toys have depicted it with as few as twelve and as many as sixteen dorsal plates, sometimes with a secondary paired row of smaller plates. It also has armor in the form of osteoderms on its body, which may be either chitinous or keratinous depending on the genes sourced to facilitate their growth.

According to the Jurassic World mobile games, including Jurassic World: The Game and Jurassic World Alive, the armor of the Stegoceratops is defensive in nature and reinforced enough to repel the attacks of Tyrannosaurus and Allosaurus. This resilience would imply that the chitinous armor derived from the rhinoceros beetle is incorporated into the plates, frill, and osteoderms of this animal.

Along with the overall body shape, the Stegoceratops demonstrates ceratopsian anatomy quite prominently in its skull. It is usually depicted with two forward-curving bovine-like horns, which the mobile game Jurassic World: The Game states are three feet in length. Originally, the game depicted it with a small nasal horn like that of Triceratops, which the sequel Jurassic World Alive and management simulator Jurassic World: Evolution also use. However, most other depictions (including the only known depiction in the hard film canon) instead depict it with only two horns and a tall upper beak or nasal boss rather than a nasal horn. If any Stegoceratops would have had a nasal horn, it would likely have been a rare genetic trait. It also has a large skull frill with sixteen small epioccipitals. It is sometimes depicted with quite large fenestrae in the frill, unlike Triceratops but much like Torosaurus, though the frill itself is shaped more like that of Triceratops. The jawline is sharp and its nostrils are flat. Some depictions show this animal with straight horns like those of Triceratops, but most opt for the bovine-like appearance which has been likened to Nasutoceratops. In Jurassic World: Evolution, its horns are extremely lengthy and slightly waved in shape, somewhat resembling Torosaurus, along with the aforementioned nasal horn and a second pair of much smaller horns next to the large ones. This version also features multiple hornlets on the frill.

Excerpt from the Jurassic World Hybrids book which states the length and weight of the Stegoceratops. However, this book features multiple discrepancies with the mobile game it used as source material and is not corroborated by any in-film evidence.

The Stegoceratops is said to be 33 feet in length in the Jurassic World Hybrids children’s book, which also describes it as weighing 11 tons. However, Jurassic World: The Game claims it weighs 15 tons. The version which appears in Jurassic World: Evolution is 9 meters (29.53 feet) long and 3 meters (9.84 feet) tall. Weight estimates for this variant have not yet been found.

Coloration in this animal is unknown, as none have appeared in the flesh. Concept art shows a forest green body with some slightly more vibrant markings, with beige coloration on the dorsal plates, fenestrae, snout, and underbelly. The small eyes appear orange or yellow. Alternate coloration depicted for this animal include dark blue with beige markings, light blue with darker blue stripes and streaks, dark gray with yellow striping, or dull green with orange outlining on some stripes. It is much more vibrantly colored in Jurassic World Alive, which features a yellow-green animal with darker splotching on its upper body and bright red plates and frill. Jurassic World: Evolution colors it similarly to the concept art, with more brightly colored plates including a dark brown streak across each.

Growth
Concept art of a mother and child Stegoceratops by Ian Joyner

In its main constituent genera, Stegosaurus and Triceratops, the hatchlings lack the bodily ornamentation that defines the adults, which grow out over time. In Triceratops, the horns and other bony skull features start out as nubs which gradually extend, curve, and sharpen with age. In Stegosaurus, the plates and thagomizer spikes are initially smaller and more rounded, achieving their adult shape and size during adolescence. Concept art of Stegoceratops featuring an infant has suggested that a similar ontogeny would have occurred in this artificial genus.

Sexual Dimorphism

There is no sexual dimorphism known for Stegoceratops; the sex of the animal rendered on Dr. Wu’s computer is not identified. Minimal sexual dimorphism is seen in Stegosaurus and Triceratops.

Habitat
Preferred Habitat

Due to a lack of observable specimens, the ideal habitat of Stegoceratops is not known. Its progenitor species have been depicted in both grassland and forest habitat; in particular, Stegosaurus has been shown both in forests and grassland, while Triceratops appears to strongly prefer grassland.

Stegoceratops depicted in a forested habitat

The abilities of Stegoceratops theoretically include camouflage, bioluminescence, and oxygen extraction from water; these would enable it to thrive in environments with considerable amounts of cover, as well as in wetland or possibly aquatic habitats.

Muertes Archipelago

There is no evidence of Stegoceratops being bred in the Muertes Archipelago. Its creator, Henry Wu, went into hiding in late 2015 before any specimens were created. At the time, Isla Sorna and the other islands of the archipelago were supposedly abandoned by Masrani Global Corporation and kept strictly under watch and off-limits to civilians.

Isla Nublar

In an early concept for Jurassic World, at least one Stegoceratops would have been encountered in Sector 5 on Isla Nublar during the December 22, 2015 incident. However, it was cut from the plot relatively early on, and in the final version no specimens had yet been created.

Mantah Corp Island

This secretive project does not seem to have gotten the attention of InGen’s rival Mantah Corporation, which operated a clandestine animal testing facility on Mantah Corp Island in the 2000s and 2010s. They did succeed at creating their own part-ceratopsian hybrid genus, but Stegoceratops escaped their knowledge.

BioSyn Genetics Sanctuary

With no confirmed wild specimens, Stegoceratops was not acquired by BioSyn Genetics after being authorized by multiple governments to capture escaped de-extinct animals. The ultimate fate of the DNA samples belonging to this species has yet to be determined; if BioSyn managed to obtain any, they may have been housed at the BioSyn Genetics Sanctuary where the company was headquartered.

Black market

Stegoceratops existed only as a genome at the time Jurassic World closed in 2015, with no known live specimens. The samples were evacuated off of Isla Nublar alongside Henry Wu when the park was shut down, removing them from InGen lab facilities and into less traceable locations. Some may have been seized by the U.S. government during raids on labs used by Wu, but this is unconfirmed. Their last known home was the Lockwood estate, where Henry Wu was in hiding between December 2015 and June 2018. After that, the fate of the Stegoceratops samples is not known.

It is unlikely that Wu would sell his prized hybrid genomes, especially before he was ready to market live specimens. If this hybrid’s DNA has entered the global black market, it was without Wu’s consent. No assets have yet turned up anywhere despite years having gone by; if the samples were not destroyed in the Lockwood Manor fire, they are in possession of an unknown party somewhere in the world.

Wild populations

The current whereabouts of any remaining Stegoceratops DNA samples are unknown. They may be in possession of BioSyn (the last employer of Henry Wu) or the United States federal government. Alternatively, they may have been acquired by an unknown person or group of people between 2015 and the present day. Finally, and sadly most likely, the samples may have been destroyed in the fire at Lockwood Manor on June 24, 2018.

Behavior and Ecology
Activity Patterns

The activity patterns of Stegoceratops are unknown. However, both of its primary constituent genetic sources are diurnal.

Diet and Feeding Behavior

Both Stegosaurus and Triceratops are herbivorous, feeding mainly on low-growing plants such as ferns and cycads. Unless features of its digestive system were altered, the Stegoceratops would most likely feed from the same kinds of plants. Its large beak would allow it to bite through even relatively tough plant matter.

In Jurassic World: Evolution, it prefers to eat rotten wood, but will also happily feed on horsetails and palms. On the other hand, pawpaws, mosses, and cycads are unhealthy for it to consume. Bracken ferns constitute a part of its diet, but in larger quantities they can cause gastric poisoning.

Social Behavior
Stegoceratops seen in profile

While the behavioral patterns of an unrealized hybrid animal cannot be predicted with certainty, both Stegosaurus and Triceratops have been shown to enjoy the company of others of their kind and are highly protective of their young. It is possible that Stegoceratops would display similar social structure to these dinosaurs.

In Jurassic World: Evolution, this animal forms herds of three to eight individuals, becoming stressed if it is lonely or overcrowded.

Reproduction

As a dinosaur, the Stegoceratops would have laid eggs. Beyond this, its reproductive behaviors are entirely speculative. Mating behaviors are not known, though Triceratops is confirmed to have a cloaca in Jurassic Park: The Game. As most of the body of Stegoceratops is anatomically similar to ceratopsians, it can be hypothesized that the Stegoceratops would also have had a cloaca.

The horns and frill of Triceratops and other ceratopsians are frequently used in courtship displays and mating contests, so Stegoceratops probably would have behaved similarly. Its dorsal plates may also have served courtship functions.

Most medium to larger herbivorous dinosaurs lay round eggs, incubating them for around six months; Stegoceratops probably incubated for a similar period of time.

Communication

Since no Stegoceratops have been observed in the flesh, its vocalizations are unknown. Its main constituent genetic sources, Stegosaurus and Triceratops, communicate using various low-pitched calls; therefore, if it were to make similar noises, Stegoceratops could theoretically vocalize a range of moans, mooing sounds, and bull-like bellows. It most likely would also use its ornate skull and dorsal plates as visual signals.

The mobile games based on Jurassic World generally give it the same sound range as Triceratops and other ceratopsids. Jurassic World: Evolution gives it a unique range of vocalizations, which include cattle-like lowing and groaning sounds.

Ecological Interactions

Thus far, no Stegoceratops have been observed in the wild, or alive at all. It would most likely feed on low-growing plants. Its horns and thagomizer would provide a good defense against most predatory animals, so most smaller carnivores would probably not threaten it.

According to the game Jurassic World: Evolution, the Stegoceratops is not affected by the common cold in any way.

Cultural Significance
Symbolism

To its creator Dr. Henry Wu, the Stegoceratops was a major achievement and a step forward in his research into the artificial hybridization of different animal genera. While his most notable successes were hybrids of theropod dinosaurs, the Stegoceratops instead consisted primarily of ornithischian traits, with its two main constituent species belonging to the clade Genasauria. Wu viewed his creations as a form of art rather than just scientific achievement, and was quite proud of the Stegoceratops in particular. He refers to it as a “queen” in Jurassic World: Evolution.

Because it is not known to the wider public, it has not yet been assigned any symbolic values by humans. Its obviously hybrid appearance would easily make it representative of this field of science. There is a minor indication in the Jurassic World Employee Handbook that it was at one point intended by InGen to represent the hybrid concept, as images of a Stegosaurus and a Triceratops appear on a page where Claire Dearing is discussing unspecified hybrid animals.

In Captivity

Since no living Stegoceratops are confirmed to have been created yet, nothing is known about exactly how to care for one in captivity. For the time being, speculation is all that is possible, based on what is known of stegosaurid and ceratopsid needs.

Science

The Stegoceratops is presumably not known to the wider scientific community at this point in time. It was only known to its creator Henry Wu and his higher-ranking genetics staff at InGen, and possibly some of his colleagues at the Lockwood laboratory after leaving the company. As of such, this dinosaur has not yet yielded any publicly-accessible scientific knowledge, leaving its potential untapped. It was a notable success in Henry Wu’s research into the hybridization of different animal genera, but no living specimens have yet been reported.

Politics

While there are no reports of living Stegoceratops so far, the other animal hybrids (the chimerid lineage, beginning with Indominus rex) created by Dr. Wu have been specialized for combat and were intended for military usage. While there is no evidence that Stegoceratops was intended specifically to be a combat animal, the concept of military bioengineering is now a precedent in the field of genetic hybridization. It certainly takes the best of its progenitors’ defensive capabilities.

Were the Stegoceratops to be bred and marketed as a military animal, it would be the first herbivorous hybrid species to be put to use in this way. It would ultimately be subject to the same controversies as the predatory hybrids; these include animal welfare, the ethics of genetically engineering new species, and the murky legality of biological weapons.

Resources

At the moment it is unknown whether Stegoceratops was originally funded specifically for use as a military animal or if it was designed with purely scientific pursuits in mind. There is minimal evidence (present in the Jurassic World Employee Handbook, but only indirectly) that it might have been intended as a Jurassic World park attraction, possibly as an alternative to the Indominus that was considered at some point.

Other hybrid species of dinosaurs have been thoroughly exploited by their creators, since all were meant to be used essentially as living weapons. Stegoceratops might face a similar fate if it is ever bred.

Safety

It is not possible to give detailed safety guidelines for a species that has never existed, but judging by its physiological traits and genetic ancestry, the behaviors of Stegoceratops can be loosely predicted. Should any be created, they should probably be treated with the same types of cautions as Stegosaurus and Triceratops.

Behind the Scenes
Stegoceratops is depicted in the Jurassic World Hybrids book as a genetically modified Stegosaurus, seen here alongside a genetically modified Triceratops. This is heavily derived from Ludia’s Jurassic World: The Game and is not corroborated by in-film evidence.

Stegoceratops would have appeared in Jurassic World during a scene in Sector 5, where it would have revealed that Henry Wu was engineering more hybrids than the Indominus. It was removed from the script at the behest of director Colin Trevorrow’s son, who suggested that the existence of other hybrids would make Indominus rex appear less unique.